This project will fill an important gap in the study of adaptive radiation through the development of analytical tools for identifying and analyzing convergence among whole radiations. I will develop a quantitative measure of convergence among independently diversifying lineages and build alternative models for convergence arising from similar adaptive landscapes, developmental constraints, or variation in the rate of evolution. I will then use these methods to investigate convergent patterns of phenotypic evolution in a classic case of replicated adaptive radiation – the Anolis lizards of the Greater Antilles.